What is the conservation status of common wombats?
The bare-nosed wombat or common wombat (Vombatus ursinus) is not listed as rare, vulnerable or endangered under the Tasmanian Threatened Species Protection Act 1995 or the Australian Environment Protection and Biodiversity Conservation Act 1999, and are listed as Least Concern by the International Union for Conservation of Nature (IUCN).
Wombats are protected in Tasmania under the Tasmanian Nature Conservation Act 2002 and permits are required to take and possess this species.
There are three subspecies of common wombat:
• Bare-nosed wombat, Vombatus ursinus hirsutus, which is found on the south-eastern Australian mainland
• Tasmanian Wombat, Vombatus ursinus tasmanicus, which is found in Tasmania, and
• Flinders Island Wombat, Vombatus ursinus ursinus, which was once found throughout the Bass Strait islands but is now restricted to Flinders Island and has been introduced to Maria Island.
Are wombat numbers declining?
NRE Tas has been monitoring counts of mammals along spotlight survey transects in eastern, northern, southern and central Tasmania for several decades. Wombat counts have generally increased across these regions since 1985. The reasons for this increase are not clear but may relate to changes in land use and management practices that increase food resources for wombats.
NRE Tas Spotlight Survey counts and monitoring by the University of Tasmania have shown that the population of wombats in the area immediately to the west of the Tamar River and in Narawntapu National Park decreased markedly between 2010 and 2016. The University of Tasmania found that the wombat population in Narawntapu National Park decreased by 94% during that time. The cause of this decline has been attributed to a severe outbreak of sarcoptic mange. A similar population decrease (80%) associated with sarcoptic mange has also been recorded on a private property in the Central Highlands of Tasmania. Local population decreases associated with mange have also been reported from mainland Australia. Wombats continue to occur at both locations where decreases have been recorded in Tasmania but at much reduced numbers.
Although mange has been present in many parts of Tasmania for over 100 years, it has not resulted in widespread declines in numbers of wombats. Understanding why outbreaks of mange occur in some areas and cause significant losses of wombats is a topic of ongoing research.
For an overview of wombat population trends see the document below:
What is mange?
Sarcoptic mange is a skin infection in mammals that is caused by a burrowing parasitic mite, Sarcoptes scabiei. It affects more than 100 mammalian species worldwide, including humans and dogs. The disease is referred to as scabies in humans and mange in other species. Australian native mammals that are known to be affected by mange include ringtail possums, brown bandicoots, koalas and common wombats.
Mange infection in an animal can result in aggressive scratching, hair loss, skin thickening and crusting, skin discoloration, open wounds (from scratching), weight loss, and in severe cases, death (as a result of secondary infection and suppressed immune system).
Where did mange originally come from and how did it get here?
It is thought that humans were the original host of sarcoptic mites, who then passed it on to their domestic animals, with subsequent spill-over events to wildlife. Current evidence strongly indicates that mange was introduced to Australia (and Tasmania) by Europeans and their domestic animals about 200 years ago. The current distribution of mange disease is considered worldwide.
Why are wombats more prone to mange and does it kill them?
Of the native Australian mammal species known to be affected by mange, wombats appear to be most impacted. Wombat burrows are believed to have good conditions for the survival of mites and the transfer of mites between wombats. Laboratory studies have shown that cool, humid conditions lead to the longest survival of mites away from their host — such conditions can occur in wombat burrows.
The disease generally occurs at low prevalence in wombat populations, but higher rates of incidence and impact can occur within localised populations. These localised outbreaks may be associated with high wombat densities, during periods of drought or nutritional stress. There has been very little research on the survivorship of wombats with mange, but current knowledge indicates that wombats with mange will eventually die if untreated as a result of secondary infections. Within most wombat populations there are wombats with and without mange suggesting that some wombats are able to resist or overcome infection. More research is required to understand the factors that result in wombats becoming infected with mange.
Can mange be eradicated?
There is no method to eradicate mange from the wild, however individual wombats and other animals can be treated for mange.
While treatment may be relatively straightforward for tame or captive animals, it is more challenging to treat animals in the wild, especially for wombats which are typically nocturnal, live underground and are not well-suited to captivity.
Preliminary
results from recent treatment trials are promising and further research is underway to
determine the best and safest way to treat wombats in the wild. Whilst treatment methods will help combat mange outbreaks and treat
individual animals, it will not eliminate mange from wild populations.
Where does mange occur?
Wombat populations throughout south-eastern Australia are affected by mange. In Tasmania, mange has been reported from northern, eastern, central and southern areas of Tasmania, including Flinders Island. No confirmed observations of mange have been recorded from western Tasmania nor on Maria Island.
The disease generally occurs at low prevalence, but more extreme outbreaks can occur within populations. It is not known why these outbreaks occur, but they appear to be associated with times of nutritional stress and/or overcrowding (i.e. high-density wombat populations).
Cool, humid conditions, which can be found in wombat burrows, support the extended survival of mites away from their hosts. Research by the University of Tasmania suggest that mite survival in burrows is longer during winter than summer.
How does mange spread?
In wombats, most transmissions are thought to occur becuase of burrow sharing. While adult wombats are mainly solitary when above ground, their home ranges overlap with other wombats and burrows can be used by more than one wombat, often at separate times but occasionally simultaneously.
Sarcoptes mites can persist in the burrows in the absence of wombats for up to three weeks. This persistence by the mites provides a reservoir for the disease to spread to other wombats.
Other mammal species entering infected wombat burrows could also spread the disease.
Is mange spreading and how widespread is mange in wombat populations?
Sarcoptic mange is extensively distributed in the bare-nosed wombat’s geographic range, including Tasmania. Of 24 locations surveyed since 2016, mange has been detected in wombats in eight, or 33%, of locations. Mange is widespread in Tasmania and has been for a very long time, but to date, no confirmed observations of mange have been recorded from western Tasmania nor on Maria Island.
In general, prevalence of mange within wombat populations is often low with little or no impact on their population numbers. However, in some localised areas where prevalence is high and outbreaks occur, such as Narawntapu National Park and surrounding areas, the impacts can be severe.
Monitoring of wombats in Tasmania is being undertaken by NRE Tas to better understand the extent of mange within wombat populations across the State.
Can mange-infected burrows be fumigated?
There are currently a number of practical challenges associated with burrow fumigation that make this technique not yet feasible. Fumigating burrows could result in exposure and adverse animal welfare outcomes for wombats that may occupy fumigated burrows. It is unknown if fumigants would sufficiently penetrate the environments in burrows that mites occupy (such as bedding chambers).
There is currently no simple technique to sample mange mites from burrows to test if fumigation is successful. Burrows can have multiple entrances and this may impact capacity to successfully fumigate a burrow system to eliminate the mange mite. Where wombats occur, there are often numerous burrows scattered across the landscape, making it difficult to determine which burrows should be fumigated and it is expensive to consider fumigating all of them.
Will mange cause wombats to become extinct?
There is no evidence to suggest that mange will cause common wombats to go extinct. Mange has been present in Australia for over 200 years. Although mange has caused severe local population declines in Australia, it has not led to a widespread reduction in the numbers of common wombats. There have been published reports of periodic local population declines of wombats since the 1930s on the Australian mainland. In Tasmania, monitoring of common wombats over more than 30 years suggests that numbers have generally increased despite decreases in the Central North and on a property in the Central Higlands.
The Tasmanian wombat population will continue to be monitored to detect signs of population change.
How do I know if a wombat has mange?
Signs of mange infection include itching, scratching, skin thickening and crusting, loss of hair and body condition. Mange infections usually present as crusty skin lesions on the sides and legs of wombats. Scabs can also form around the eyes and ears, impacting on the animal’s sight and hearing.
However, not all skin conditions on wombats can be attributed to mange. Wombats may be affected by other diseases that cause some of the symptoms of mange. Wombats may become aggressive towards each other resulting in wounds. These attack wounds, which may look like mange in appearance, are usually on the wombat’s rump area whereas the back and rump are usually the least affected areas on a wombat with mange.
Wombat showing severe condition resulting from mange
copyright: K. Simpson
How long does it take for the infection to spread?Typically, the first clinical signs of mange develop 1–3 weeks after exposure to the parasites; more chronic signs appear after 4–5 weeks. In severe cases, death may occur as a result of secondary bacterial infection 2–3 months after infection. The severity of infection may vary with initial doses of exposure as well as the condition and health status of the host.
Why are some wombats severely affected and not others?
The reasons why some wombats become infected and others do not are poorly known. Sarcoptic mange occurs in common wombat populations across most of their range, but the disease varies widely in its expression and impacts on local populations. Wombat densities, environmental conditions and wombat health are all thought to have an influence on whether wombats become infected with mange.
It is commonly reported that increased rates of mange disease in wombats are often associated with high wombat densities, and periods of environmental stress, e.g. drought and winter conditions. Expression of disease may be seasonal, resulting from increased mortality of mites due to adverse conditions (hot and dry). The overall health and condition of individual wombats also varies, and the disease may affect individual wombats that are already compromised compared to healthy robust individuals.
Recent research in Tasmania by UTAS and NRE Tas found that at small spatial scales, prevalence of mange varies spatially (3.1–37.5%), with the probability of disease greater in wombats with minimal vegetation and with high water tables in their home range, as well as where there were more burrows available per wombat. These results suggest that individuals inhabiting less optimal habitat (pasture) may be at greater risk of disease, or that diseased wombats may be competitively excluded from more optimal habitat (vegetated areas).
Does mange affect the behaviour of wombats?
Yes, mange affected wombats are more active outside of their burrows for more extended periods compared to healthy wombats and spend more time above ground during daylight hours. Mange-affected wombats spend less time walking, and more time scratching, foraging and drinking than healthy wombats. They have a slower feeding rate and they are generally in poorer body condition. As a result of scabs around the face in more severe cases, the wombats may appear deaf and / or blind, and may not be very alert or responsive.
When are infected wombats observed most often?
Wombats with mange can be observed at any time of day; however, an Australian mainland study and the results from studies in Tasmania found that wombats with mange are more likely to be observed during daylight hours than healthy wombats. A possible reason for this change in behaviour is likely to be that mange-affected wombats have a higher energy requirement necessitating the continuous search for food. The increased daylight activity by mange-affected wombats may also be related to the vision impairment of severely mange-infected wombats. Whatever the cause of this behavioural change, it is possible to get a misleading impression of mange prevalence from daylight observations.
Day time versus nighttime observations
Click image to enlarge - The ratio of healthy wombats to mange-affected wombats observed in the night and day varies because healthy wombats typically feed at night. Mange-affected wombat numbers remained steady both in the day and at night.
How can mange be treated?
Individual wombats and other animals can be treated for mange. While this may be relatively straight forward for tame or captive animals, it is more challenging to treat animals in the wild, especially for wombats which are typically nocturnal, live underground and are not well suited to captivity.
The current treatment of mange in Wombats requires regular doses of moxidectin (Cydectin) using the method of burrow flaps or pole and scoop. The number and recurrance of treatment depends on the dosage amount (high or low dose). Authorisation from the Australian Pesticides and Veterinary Medicines Authority is required, and permits may be required from NRE Tas depending on the method of treatment administration.
Recommended treatment methods are delivery of the acaricides (pesticides that kill mites and ticks) via a burrow flap over a wombat burrow that doses the wombat as it enters and exits the burrow, or by direct application straight onto the wombat via a scoop on a pole. The pole and scoop method is generally used for badly affected wombats that can be easily approached. Wombats should not be chased to apply treatment.
Two studies of the effectiveness of treating mange in common wombats using the burrow flap method have been conducted in the wild, one in Tasmania and one in NSW. Results indicate that the effectiveness of the treatment regime is variable at the population level. Both studies highlighted the considerable logistical challenges in treating all individuals and all burrows.
Multiple doses of Cydectin are required, as the drug isn’t easily absorbed through affected skin and therapeutic drug levels need to be maintained for long enough for the skin condition to improve and the animal to resist re-infection from the environment.
A new treatment method, using Fluralaner (Bravecto) has now been approved for use on wombats by the Australian Pesticides and Veterinary Medicines Authority. Treatment involves a spot-on dose that is applied to the back of the wombat using a pipette. Persons who are trained and experienced in the use of veterinary chemicals, who have appropriate experience in identifying mange infection and fluralaner toxicity signs in wombats may use this product. Permits may also be required from NRE Tas depending on the method of treatment administration.
Reinfection of treated wombats can occur if infected wombats remain in the population, if other hosts transmit mites to wombats, or if mites remain viable and infectious in the landscape. Due to the potential for the mites to develop resistance to the acaricides continuous treatment is not encouraged.
Wombat exiting burrow fitted with a burrow flap
copyright: A. Martin
Should wombats be treated in captivity or in the wild?
Wherever possible wombats should be treated for mange in the wild. Holding a wild wombat in captivity comes at some risk to the wombat due to stress associated with capture, handling and confinement. This stress is exacerbated in mange affected wombats whose immune systems are already compromised.
The survival of captive-raised or captive-held wombats post-release is not well understood and evidence from a limited number of captive-release studies on other animals suggests that survival may be poor. In some circumstances, bringing wombats into captivity to treat mange may be appropriate provided suitable facilities are available and that the animals will be released an area close to where they originated.
Does mange occur in other species?
Mange has been reported in over 100 species of mammal, including humans. The disease is referred to as scabies in humans and mange in other species. The broad range of hosts for sarcoptic mange includes domestic dogs, livestock (e.g. cattle, pigs, goats, camels) and wildlife (e.g. red foxes, coyotes, wolves, deer, bobcats, wombats, koalas and wallabies).
Why don’t wallabies get mange as badly as wombats?
Mange has been recorded in some species of wallabies. Wallabies do not get exposed to the mites that cause mange as often as wombats because of their different social behaviour and resting places. Wombat burrows are believed to be good places for the mites to survive and for transmission between wombats.
Can Tasmanian Devils contract mange?
Devils sometimes show mild symptoms of mange, but the causal agent is a devil-specific mite, Satanicoptes armatus, not Sarcoptes scabiei. Mange due to Sarcoptes scabiei has not been recorded in Tasmanian devils. Sarcoptes scabiei mites can infect a range of different mammalian hosts, with clinical severity varying among species. Wombats, dogs and foxes are particularly susceptible to developing mange symptoms, especially if they have a weakened immune system that makes them more vulnerable to infections.
What problems do wombats cause for landholders?
The main concern landholders raise about wombats is the damage they can cause to fences, especially wallaby proof fences, allowing wallabies access to paddocks resulting in damage to pastures and crops. Wombats can contribute to pasture loss, they can also damage dam walls and other infrastructure, and their burrows can be a hazard for livestock, as well as for horse and motorbike riders.
How can landholders deal with wombats impacting their properties?
Landholders are encouraged to adopt non-lethal approaches to managing impacts of wombats on their properties such as installing wombat gates to allow wombats, but not wallabies, to enter and exit pasture areas. However, NRE Tas recognises that these approaches are not always possible or successful in managing the impacts.
Wombats are protected wildlife and a permit is required to use lethal methods. Applications for a permit to take wombats are assessed on a case-by-case basis, which includes a visit to the applicant’s property. A range of factors are taken into account including the local abundance of wombats, the impacts that the wombats may be having on the property, and whether or not non-lethal measures will be effective at the site. If a permit is issued, it will specify the number of wombats that are authorised to be taken and the timeframe during which the permit applies.
There is currently a moratorium on the issuing of property protection permits in the West Tamar region due to the decrease in the wombat population in the area.
What happened to the wombats at Narawntapu National Park?
Wombats were previously a common species in Narawntapu National Park in central northern Tasmania. Although mange has been present in the park for decades, there was an outbreak of mange in 2006 following a severe drought. Since then, mange has resulted in a substantial reduction in wombat numbers. Monitoring by the University of Tasmania between 2010 and 2016 shows that the wombat population declined by 94%. On-going annual monitoring by NRE Tas shows that wombats continue to occur in the park, but at very low numbers.
What was done to help the wombats in the Narawntapu National Park?
The University of Tasmania, with assistance from the Parks and Wildlife Service, undertook an experiment to evaluate if mange could be eliminated from the wombat population that remained in the western end of Narawntapu National Park.
This involved attempting to treat the entire wombat population with burrow flaps and Cydectin. This was a labour-intensive program that involved deploying and maintaining >200 burrow flaps on all known wombat burrows that were active.
Although the program was temporarily successful in treating some individuals in the population, it failed to treat all wombats and thus eliminate the disease. This research showed that it is very difficult to ensure that all wombat burrows and burrow entrances are found and that all wombats will receive the full treatment regime. While treating individual wombats can be successful, treating whole populations is very difficult. Another consideration is that, even when individual wombats have been treated and recover, they are susceptible to reinfection.
NRE Tas is monitoring the wombat population at Narawntapu to inform management options.
Will Narawntapu National Park ever have wombats again?
Ongoing annual wombat monitoring by NRE Tas shows that a small number of wombats still occur in Narawntapu National Park. It is not known how long it will take for the population to recover. There is potential for wombats to self-introduce from surrounding areas.
If a new wombat population establishes in Narawntapu National Park will it be re-infested with mange?
As mites live on a range of host species, there can be no guarantees that a new wombat population won’t be re-infested.
Is mange the only factor that impacts on wombats in Tasmania?
No, whilst mange is the
most significant factor affecting their health, there are other factors that
impact wombats. Similar to a range of native animal species, wombats are susceptible
to roadkill. Drivers can play an important role in reducing this threat by
being aware of wildlife hotspots and reducing speed at key times between dusk
and dawn. Road killed wombats can be reported via the Tasmanian Roadkill Reporter App.
Uncontrolled dogs can also impact on wombats which is why it is important to ensure that dogs remain under effective control and are not allowed to roam unsupervised.
Road-kill wombat
copyright: NRE Tas
How can I help?
There are many ways you can help:
Report observations of injured wombats or wombats with mange to NRE Tas by calling 03 61654305 or via email Wildlife.Services@nre.tas.gov.au.
Treat mange affected wombats in your area – contact NRE Tas;
Drive carefully in areas where wombats are present to reduce the number of wombats killed or injured by cars;
Report road-killed wombats (and other wildlife) via the Tasmanian Roadkill Reported App.
Use non-lethal methods to manage wombats on agricultural land, including “wombat gates” to allow their passage through fences; and
Prevent dogs from roaming in areas where wombats occur;
If you would like information material about mange to display and hand out we have the following resources available;
What do I do if I see a mange-infected wombat?
Report to NRE Tas by calling 03 61654305 or via email Wildlife.Services@nre.tas.gov.au to provide details of the location and condition of the individual.
Photographs are very useful to help assess if a wombat has mange and the severity of the mange.
Do I need permits to treat mange-infected wombats in Tasmania?
Yes, permits are required to treat mange in wombats, both in the wild and in captivity.
If you would like to treat a mange-affected wombat in your area, please contact NRE Tas for advice before commencing treatment.
NRE Tas staff can then assess:
Whether the wombat you want to treat has mange or not - providing images of the wombat is very helpful to assess this
An appropriate treatment method
Whether or not the wombat is already being responded to
What permits will be required and how to obtain permits
How to contact experienced treaters
If you have experience in treating wombats for mange and wish to apply for a permit to treat Wombats with mange, please downland the following application form:
Sulphur compounds, including lime sulphur and yellow powdered sulphur are not considered suitable as a treatment for mange in wombats on veterinary advice.
Lime sulphur:
There is some evidence of lime sulphur's ability to treat mange in dogs and cats but the compound is toxic and caustic at higher concentrations forming hydrogen sulphide gas.
The substance is likely to be an irritant and painful to skin already inflamed by mange.
The process of using sulphur compounds would require capturing, restraining, wetting and scrubbing the animal weekly for up to six weeks which would be extremely stressful for a wild animal. There is a high risk of splashes into eyes or mucus membranes, or aspiration into the lungs, more likely in a wild animal that may be struggling and requires physical restraint.
Yellow powdered sulphur:
Yellow powdered sulphur in ointment form, or mixed with various oils and applied topically, has some evidence of treating against sarcoptic mange in a variety of animal species.
Powdered sulphur is not as caustic as lime sulphur. Recommended treatment frequency ranges from twice daily to every three days for a period of weeks. The requirement for regular restraint and topical application to a large part of the body render this treatment less suitable for a wild animal than other available therapies. It has been found to be less effective, and require more treatments, than ivermectin and related drugs.
A topical treatment of Cydectin is the recommended treatment as it is safer, less stressful for the animal, and more effective than use of sulphur compounds.
Information sources and further reading
Browne E, Driessen MM, Ross R, Roach M & Carver S. (2021). Environmental suitability of bare-nosed wombat burrows for Sarcoptes scabiei. International Journal for Parasitology: Parasites and Wildlife 16, 37–47.
Burgess LG, Richards SA, Driessen MM, Wilkinson V, Amin RJ & Carver S. (2023). Fine-Scale Landscape Epidemiology: Sarcoptic Mange in Bare-Nosed Wombats (Vombatus ursinus). Transboundary and Emerging Diseases. Pages https://doi.org/10.1155/2023/2955321.
Beeton NJ, Carver S, & Forbes LK (2019). A model for the treatment of environmentally transmitted sarcoptic mange in bare-nosed wombats (Vombatus ursinus). Journal of Theoretical Biology 462, 466-474.
Borchard P, Eldridge DJ & Wright IA (2012) Sarcoptes mange (Sarcoptes scabiei) increases diurnal activity of bare-nosed wombats (Vombatus ursinus) in an agricultural riparian environment’ Mammalian Biology 77, 244-248
Carver S, Stannard GL & Martin AM (2024). The distinctive biology and characteristics of the bare-nosed wombat (Vombatus ursinus). Annual Review of Animal Biosciences 12, 135-160.
Carver S, Charleston M, Hocking GJ, Gales R & Driessen MM (2021). Long-term Spatiotemporal Dynamics and Factors Associated with Trends in Bare-Nosed Wombats. Journal of Wildlife Management 83, 449–461.
Driessen MM, Gales R, Hehn K, Dewar E, & Dobner G. (2020). Wombat gates effectively exclude browsing mammals from pasture and allow passage of common wombats. Australian Mammalogy 42, 375–379.
Driessen MM, Dewar E, Carver & Gales R (2022). Conservation status of common wombats in Tasmania I: incidence of mange and its significance. Pacific Conservation Biology 28, 103-114.
Driessen MM, Dewar E, Carver S, Lawrence C & Gales R (2022). Conservation status of common wombats in Tasmania II: population distribution and trends, and the incidence and significance of roadkill. Pacific Conservation Biology 28, 115-123.
Fraser TA, Charleston M, Martin A, Polkinghorne A & Carver S. (2016). The emergence of sarcoptic mange in Australian wildlife: an unresolved debate. Parasite and Vectors 9, 316.
Gray DF (1937) Sarcoptic mange affecting wild fauna in New South Wales. Australian Veterinary Journal 13, 154-155
Knoblauch W Carver S Driessen MM Gales R & Richards SA (2023). Abundance and population growth estimates for bare-nosed wombats. Ecology and Evolution 13.
Martin RW, Handasyde KA & LF Skerratt (1998). Current distribution of sarcoptic mange in wombats. Australian Veterinary Journal, 76: 411-414
Martin AM, Burridge CP, Ingram J, Fraser TA, and Carver S (2018). Invasive pathogen drives host population collapse: Effects of a travelling wave of sarcoptic mange on bare-nosed wombats. Journal of Applied Ecology 55, 331–341.
Martin AM, TA Fraser, Lesku JA, Simpson K, Roberts GL, Garvey J, Polkinghorne A, Burridge CP & Carver S (2018). The cascading pathogenic consequences of Sarcoptes scabiei infection that manifest in host disease. Royal Society Open Science 5: 1–14.
Martin AM, Ricardo H, Tompros A, Fraser TA, Polkinghorne A & Carver S (2019). Burrows with resources have greater visitation and may enhance mange transmission among wombats. Australian Mammalogy 41, 287-290.
Martin AM, Richards SA, Fraser TA, Polkinghorne A, Burridge CP & Carver (2019). Population-scale treatment informs solutions for control of environmentally transmitted wildlife disease. Journal of Applied Ecology 56, 2363-2375.
Ringwaldt EM, Brook BW, Buettel JC, Cunningham CX, Fuller C, Gardiner R, Hamer R, Jones M, Martin AM & Carver S (2023). Host, environment, and anthropogenic factors drive landscape dynamics of an environmentally transmitted pathogen: Sarcoptic mange in the bare-nosed wombat. Journal of Animal Ecology 92, 1786-1801.
Simpson, K, Johnson CN & Carver S (2016). ‘Sarcoptes scabiei: The mange mite with mighty effects on the common wombat (Vombatus ursinus). PLoS ONE 11(3), 1–17.
Skerratt LF (2003). Clinical response of captive common wombats (Vombatus ursinus) infected with Sarcoptes scabiei var. wombati.’ Journal of Wildlife Disease 39, 179-192
Skerratt LF (2005). Sarcoptes scabiei: an important exotic pathogen of wombats’ Microbiology Australia 26, 79-81.
Skerratt LF, Martin RW & Handasyde KA (1998). Sarcoptic mange in wombats. Australian Veterinary Journal 76, 408-410
Skerratt LF, Skerratt JH, Banks S, Martin R & Handasyde KA (2004). Aspects of the ecology of common wombats (Vombatus ursinus), Australian Journal of Zoology, 52, 303-330.